RT journal article T1 A Spanish dancer? No! A troupe of dancers: a review of the family Hexabranchidae Bergh, 1891 (Gastropoda, Heterobranchia, Nudibranchia): A troupe of Spanish dancer A1 Junqueira de Azevedo Tibirica, Yara A1 Pola Perez, Marta A1 Pittman, Cory A1 Gosliner, Terrence M. A1 Malaquias, Manuel Antonio E. A1 Cervera Currado, Juan Lucas A2 Biología K1 Hexabranchus K1 Integrative taxonomy K1 Nudibranchs K1 Ontogeny K1 Phylogeny K1 Species delimitation AB Color ontogeny and variations associated with discrete morphological differences may generate taxonomical challenges, which requires multiple data types and in-depth historical review. The nudibranch known as the Spanish dancer, Hexabranchus sanguineus, is a classic example with over 200 years of taxonomic confusion. Currently, H. sanguineus is accepted by most authors as a single species from the Indo-Pacific Ocean with Hexabranchus morsomus as a valid species from the Atlantic Ocean. Yet, despite these species being highly studied, their systematic status remains debatable. Over 30 synonyms have been proposed for H. sanguineus and even a distinct genus for H. morsomus. Here we provide, for the first time, a comprehensive review of all proposed names and an integrative taxonomic revision of the genus including morphological and molecular data. Our results reveal that H. sanguineus is a complex of five species: four previously described and an undescribed species, one of the largest nudibranchs in the world: Hexabranchus giganteus sp. nov. The genus Caribranchus is considered a junior synonym of Hexabranchus Ehrenberg, 1828 and the ontogeny of color pattern is discussed. PB Springer SN 1439-6092 YR 2023 FD 2023-06-20 LK http://hdl.handle.net/10498/28972 UL http://hdl.handle.net/10498/28972 LA eng DS Repositorio Institucional de la Universidad de Cádiz RD 21-sep-2026